BLASTX nr result
ID: Paeonia24_contig00005979
seq
BLASTX 2.2.25 [Feb-01-2011] Reference: Altschul, Stephen F., Thomas L. Madden, Alejandro A. Schaffer, Jinghui Zhang, Zheng Zhang, Webb Miller, and David J. Lipman (1997), "Gapped BLAST and PSI-BLAST: a new generation of protein database search programs", Nucleic Acids Res. 25:3389-3402. Query= Paeonia24_contig00005979 (1425 letters) Database: ./nr 38,876,450 sequences; 13,856,398,315 total letters Searching..................................................done Score E Sequences producing significant alignments: (bits) Value emb|CBI17720.3| unnamed protein product [Vitis vinifera] 118 6e-24 ref|XP_002265001.1| PREDICTED: flavin-containing monooxygenase F... 118 6e-24 emb|CBI24792.3| unnamed protein product [Vitis vinifera] 112 4e-22 ref|XP_002265037.1| PREDICTED: flavin-containing monooxygenase F... 108 8e-21 ref|XP_003542842.1| PREDICTED: flavin-containing monooxygenase F... 102 4e-19 ref|XP_004511006.1| PREDICTED: flavin-containing monooxygenase F... 100 1e-18 ref|XP_003627756.1| Dehydrosqualene desaturase [Medicago truncat... 97 2e-17 ref|XP_007148978.1| hypothetical protein PHAVU_005G030400g [Phas... 97 2e-17 ref|XP_006591170.1| PREDICTED: flavin-containing monooxygenase F... 94 2e-16 ref|XP_006591168.1| PREDICTED: flavin-containing monooxygenase F... 94 2e-16 ref|XP_004511004.1| PREDICTED: flavin-containing monooxygenase F... 93 3e-16 ref|XP_003627754.1| Flavin-containing monooxygenase FMO GS-OX5 [... 92 4e-16 gb|EXB37532.1| Flavin-containing monooxygenase FMO GS-OX-like 3 ... 91 1e-15 gb|AFK42203.1| unknown [Lotus japonicus] 84 2e-13 gb|ACR10275.1| flavin-monooxygenase [Brassica rapa subsp. pekine... 75 5e-11 gb|AGS49162.1| flavin-monooxygenase, partial [Eruca vesicaria su... 73 4e-10 ref|XP_006417260.1| hypothetical protein EUTSA_v10007582mg [Eutr... 72 6e-10 ref|XP_006366746.1| PREDICTED: flavin-containing monooxygenase F... 71 1e-09 ref|XP_007157335.1| hypothetical protein PHAVU_002G061500g [Phas... 71 1e-09 ref|XP_006307454.1| hypothetical protein CARUB_v10009080mg [Caps... 71 1e-09 >emb|CBI17720.3| unnamed protein product [Vitis vinifera] Length = 883 Score = 118 bits (296), Expect = 6e-24 Identities = 59/123 (47%), Positives = 77/123 (62%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A VLSGKVLLPT G+PK +TH LP E +Y+N LLA Sbjct: 761 ELQSKWLAHVLSGKVLLPTEEEMMSDVENYYHHMEETGVPKSFTHVLPPNEIEYRNWLLA 820 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLLANRSQGAAGQ 362 ++ +PPL+EWR +++ + +KF K+ G DQWDDDYWDAVI S+ + LAN GQ Sbjct: 821 QVEMPPLKEWRGRMYRECVKFAKAKLDGYRDQWDDDYWDAVIASQEVNHLAN------GQ 874 Query: 363 ETG 371 + G Sbjct: 875 DVG 877 Score = 108 bits (269), Expect = 8e-21 Identities = 50/105 (47%), Positives = 66/105 (62%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A VLSGK LLPT G+PK +TH LP E +Y N LLA Sbjct: 365 ELQSKWLAHVLSGKALLPTKEEMMSDIEKYYHHMEETGVPKRFTHVLPPNEIEYMNWLLA 424 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSK 317 ++G+PPL+EW +++++ K+ + G DQWDDDYWDAVI S+ Sbjct: 425 QVGMPPLKEWTGRMYKECAMLAKAKRDGYKDQWDDDYWDAVIASQ 469 >ref|XP_002265001.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX-like 2-like [Vitis vinifera] Length = 512 Score = 118 bits (296), Expect = 6e-24 Identities = 59/123 (47%), Positives = 77/123 (62%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A VLSGKVLLPT G+PK +TH LP E +Y+N LLA Sbjct: 390 ELQSKWLAHVLSGKVLLPTEEEMMSDVENYYHHMEETGVPKSFTHVLPPNEIEYRNWLLA 449 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLLANRSQGAAGQ 362 ++ +PPL+EWR +++ + +KF K+ G DQWDDDYWDAVI S+ + LAN GQ Sbjct: 450 QVEMPPLKEWRGRMYRECVKFAKAKLDGYRDQWDDDYWDAVIASQEVNHLAN------GQ 503 Query: 363 ETG 371 + G Sbjct: 504 DVG 506 >emb|CBI24792.3| unnamed protein product [Vitis vinifera] Length = 261 Score = 112 bits (280), Expect = 4e-22 Identities = 52/105 (49%), Positives = 68/105 (64%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A VLSGKVLLPT G+PK +TH LP E +Y+N LLA Sbjct: 132 ELQSKWLAHVLSGKVLLPTEEEMMSDVENYYHHMEETGVPKSFTHVLPPNEIEYRNWLLA 191 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSK 317 ++ +PPL+EWR +++ + +KF + G DQWDDDYWDAVI S+ Sbjct: 192 QVEMPPLKEWRGRMYRECVKFANAKPDGYRDQWDDDYWDAVIASQ 236 >ref|XP_002265037.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX-like 3-like [Vitis vinifera] Length = 448 Score = 108 bits (269), Expect = 8e-21 Identities = 50/105 (47%), Positives = 66/105 (62%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A VLSGK LLPT G+PK +TH LP E +Y N LLA Sbjct: 333 ELQSKWLAHVLSGKALLPTKEEMMSDIEKYYHHMEETGVPKRFTHVLPPNEIEYMNWLLA 392 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSK 317 ++G+PPL+EW +++++ K+ + G DQWDDDYWDAVI S+ Sbjct: 393 QVGMPPLKEWTGRMYKECAMLAKAKRDGYKDQWDDDYWDAVIASQ 437 >ref|XP_003542842.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX5-like [Glycine max] Length = 439 Score = 102 bits (254), Expect = 4e-19 Identities = 49/102 (48%), Positives = 62/102 (60%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ KWVA+VLSGKVLLPT G+PK YTH+L ++ YK+ L+A Sbjct: 330 ELQCKWVAKVLSGKVLLPTEKEMMESVKEYYQLMEENGLPKRYTHSLYPLQADYKHWLVA 389 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVI 308 EIGLPPLEEW+E + + K + D+WDD YWDA+I Sbjct: 390 EIGLPPLEEWKENMLKQCFKNFVEMNEKYRDEWDDTYWDAII 431 >ref|XP_004511006.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX5-like [Cicer arietinum] Length = 438 Score = 100 bits (250), Expect = 1e-18 Identities = 46/104 (44%), Positives = 67/104 (64%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ+KWVA+VLSGKV+LPT G+PK YTH+L + +YK+ L++ Sbjct: 331 ELQAKWVAKVLSGKVVLPTEEEMMESVKELYKFMEENGLPKRYTHSLRPFQVEYKHWLVS 390 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKS 314 EIGLPPLE+WR+ + + K + + D+WDD+YWD++I+S Sbjct: 391 EIGLPPLEDWRDNMLMECYKKLIEMNEMYRDEWDDNYWDSIIQS 434 >ref|XP_003627756.1| Dehydrosqualene desaturase [Medicago truncatula] gi|355521778|gb|AET02232.1| Dehydrosqualene desaturase [Medicago truncatula] Length = 431 Score = 97.1 bits (240), Expect = 2e-17 Identities = 44/104 (42%), Positives = 62/104 (59%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ+KWVARVLSGK+LLPT +PK YTH++ + YK L A Sbjct: 324 ELQAKWVARVLSGKILLPTEEEMMESVKDMYRVMEENALPKRYTHSIRPFQADYKQWLAA 383 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKS 314 +IGLPPLE+WR+ + + K + + D+WDD+YWD +I++ Sbjct: 384 QIGLPPLEDWRDNMLMECFKKVIEMNEMYRDEWDDNYWDTIIEN 427 >ref|XP_007148978.1| hypothetical protein PHAVU_005G030400g [Phaseolus vulgaris] gi|561022242|gb|ESW20972.1| hypothetical protein PHAVU_005G030400g [Phaseolus vulgaris] Length = 437 Score = 96.7 bits (239), Expect = 2e-17 Identities = 49/105 (46%), Positives = 63/105 (60%), Gaps = 1/105 (0%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ KWVARVLSGKVLLPT G PKH TH + E Y N L A Sbjct: 330 ELQCKWVARVLSGKVLLPTEKEMLADVEEYYQQMEKDGFPKHMTHYVHFKEIGYCNWLAA 389 Query: 183 EIGLPPLEEWREKLHEDTIK-FIKSVQRGSSDQWDDDYWDAVIKS 314 E GLPP+E WRE+++ ++IK F+ + D+WDD +W+A+IK+ Sbjct: 390 EAGLPPIEHWREEMYLESIKPFLLGMLENYRDEWDDAHWNAIIKN 434 >ref|XP_006591170.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX-like 3-like isoform X3 [Glycine max] Length = 370 Score = 93.6 bits (231), Expect = 2e-16 Identities = 49/104 (47%), Positives = 60/104 (57%), Gaps = 1/104 (0%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ KWVARVLSGKVLLPT G PKH TH L E Y N L A Sbjct: 258 ELQCKWVARVLSGKVLLPTEKEMMAYVEEYYQQMEKDGFPKHMTHYLHFKEIGYCNWLAA 317 Query: 183 EIGLPPLEEWREKLHEDTIK-FIKSVQRGSSDQWDDDYWDAVIK 311 + GLPP+E WR+ ++ ++IK + +Q DQWDD +W A+IK Sbjct: 318 KAGLPPIEHWRDAMYLESIKPVLLGLQDNYRDQWDDAHWKAIIK 361 >ref|XP_006591168.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX-like 3-like isoform X1 [Glycine max] Length = 443 Score = 93.6 bits (231), Expect = 2e-16 Identities = 49/104 (47%), Positives = 60/104 (57%), Gaps = 1/104 (0%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ KWVARVLSGKVLLPT G PKH TH L E Y N L A Sbjct: 331 ELQCKWVARVLSGKVLLPTEKEMMAYVEEYYQQMEKDGFPKHMTHYLHFKEIGYCNWLAA 390 Query: 183 EIGLPPLEEWREKLHEDTIK-FIKSVQRGSSDQWDDDYWDAVIK 311 + GLPP+E WR+ ++ ++IK + +Q DQWDD +W A+IK Sbjct: 391 KAGLPPIEHWRDAMYLESIKPVLLGLQDNYRDQWDDAHWKAIIK 434 >ref|XP_004511004.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX5-like isoform X1 [Cicer arietinum] Length = 430 Score = 92.8 bits (229), Expect = 3e-16 Identities = 46/104 (44%), Positives = 62/104 (59%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ+ WVARVLSG +LLPT G+PK Y + +++YKN L A Sbjct: 323 ELQATWVARVLSGNILLPTEEEMLESVQDIYREMEKKGLPKRYYLIKFVWQSEYKNWLAA 382 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKS 314 +IGLPPLEEWRE + ++ IK + DQWDD YW+A+I++ Sbjct: 383 QIGLPPLEEWRENMLKECIKKLIETPTKYRDQWDDAYWNAIIQT 426 >ref|XP_003627754.1| Flavin-containing monooxygenase FMO GS-OX5 [Medicago truncatula] gi|355521776|gb|AET02230.1| Flavin-containing monooxygenase FMO GS-OX5 [Medicago truncatula] Length = 432 Score = 92.4 bits (228), Expect = 4e-16 Identities = 44/104 (42%), Positives = 63/104 (60%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQ+KW+ARVLSGKVLLP G+ K TH+L ++ YK+ L+ Sbjct: 325 ELQAKWIARVLSGKVLLPDEEEMMASIKDFYQSMEENGLSKRQTHSLRPLQGDYKHWLVE 384 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKS 314 +IGLPPLE+WR+ + + +K + D+WDD+YWDA+I+S Sbjct: 385 QIGLPPLEDWRDNMLMECLKNSIEMNEMFRDEWDDNYWDAIIQS 428 >gb|EXB37532.1| Flavin-containing monooxygenase FMO GS-OX-like 3 [Morus notabilis] Length = 438 Score = 91.3 bits (225), Expect = 1e-15 Identities = 45/104 (43%), Positives = 65/104 (62%), Gaps = 1/104 (0%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+ARVLSGK++LP+ G+PK TH L L + Y+N L A Sbjct: 331 ELQSKWIARVLSGKLMLPSEEEMMASTEEFYKKMEENGLPKRSTHFL-LFQIGYQNWLCA 389 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQR-GSSDQWDDDYWDAVIK 311 ++GLPP+EEW+ K++E+++ I ++ DQWDD YW+ +IK Sbjct: 390 QLGLPPIEEWKYKMYEESLNNIYEMRDIRYKDQWDDAYWEGIIK 433 >gb|AFK42203.1| unknown [Lotus japonicus] Length = 184 Score = 84.0 bits (206), Expect = 2e-13 Identities = 40/87 (45%), Positives = 55/87 (63%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKWVAR+LSGKVLLPT G+PK YTHAL + YK+ L+A Sbjct: 95 ELQSKWVARILSGKVLLPTEEGMMEDIRNFYQFMEENGLPKRYTHALRPFQVDYKDWLVA 154 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQR 263 +IGLPPLE+W E+++ + K + +++ Sbjct: 155 QIGLPPLEDWMEQMYSECFKNLVEMKK 181 >gb|ACR10275.1| flavin-monooxygenase [Brassica rapa subsp. pekinensis] Length = 459 Score = 75.5 bits (184), Expect = 5e-11 Identities = 43/114 (37%), Positives = 59/114 (51%), Gaps = 1/114 (0%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLI-ETKYKNCLL 179 ELQSKWVA VLSG+V LP+ G PK YTH L ++ + +Y N + Sbjct: 338 ELQSKWVAAVLSGRVTLPSEDKMMEDTNALYTKRDANGFPKRYTHRLGVVGQAEYLNWIA 397 Query: 180 AEIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLLANR 341 +IG PP+E+WR K ED + + D+WDDD+ +I Y D L + Sbjct: 398 DQIGEPPVEQWRNKELEDGYVRLATQADTFRDKWDDDH---LIAEAYEDFLRQK 448 >gb|AGS49162.1| flavin-monooxygenase, partial [Eruca vesicaria subsp. sativa] Length = 333 Score = 72.8 bits (177), Expect = 4e-10 Identities = 41/113 (36%), Positives = 56/113 (49%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKWVA VLSG+V LP+ G PK YTH L + +Y N + Sbjct: 213 ELQSKWVAAVLSGRVTLPSEEKMMEDTNALYAKRDANGFPKRYTHRLGASQAEYLNWIAD 272 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLLANR 341 +IG PP+E+WR + E + + D+WDDD+ +I Y D L + Sbjct: 273 QIGEPPVEQWRNQELEGGYVRLATQPDTFRDKWDDDH---LIVEAYEDFLRQK 322 >ref|XP_006417260.1| hypothetical protein EUTSA_v10007582mg [Eutrema salsugineum] gi|557095031|gb|ESQ35613.1| hypothetical protein EUTSA_v10007582mg [Eutrema salsugineum] Length = 461 Score = 72.0 bits (175), Expect = 6e-10 Identities = 39/113 (34%), Positives = 57/113 (50%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKWVA VLSG+V LP+ G+PK YTH L + +Y N + Sbjct: 338 ELQSKWVAAVLSGRVTLPSEDKMMEDVTAFYAKRDANGLPKRYTHRLGASQIEYLNWIAE 397 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLLANR 341 ++G PP+E+WR + E + + D+WDDD+ +I Y D + + Sbjct: 398 QVGAPPVEQWRGQEVEGGYFRLATQSDTFRDKWDDDH---LIVEAYEDFVRQK 447 >ref|XP_006366746.1| PREDICTED: flavin-containing monooxygenase FMO GS-OX-like 4-like isoform X1 [Solanum tuberosum] Length = 480 Score = 70.9 bits (172), Expect = 1e-09 Identities = 38/104 (36%), Positives = 58/104 (55%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A VLSG++ LP+ + +PK YTH + + Y + L A Sbjct: 360 ELQSKWIAGVLSGRISLPSKEDMNADIEAFYSSMAASCIPKRYTHNMDDSQFDYDDWLAA 419 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKS 314 E G PP EEWR++++ + K K++ D+WDDD D +I++ Sbjct: 420 ECGSPPSEEWRKQMYFISRKNKKTLPETYRDEWDDD--DLIIQA 461 >ref|XP_007157335.1| hypothetical protein PHAVU_002G061500g [Phaseolus vulgaris] gi|561030750|gb|ESW29329.1| hypothetical protein PHAVU_002G061500g [Phaseolus vulgaris] Length = 453 Score = 70.9 bits (172), Expect = 1e-09 Identities = 36/110 (32%), Positives = 54/110 (49%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKW+A +LS ++ LP+ G PK YTH + L++ Y N + Sbjct: 343 ELQSKWIAGILSNRIALPSKEEMVEDIEVFYSSLEATGTPKRYTHNMALLQWDYNNWIAD 402 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLL 332 + GLP EEWR++++ T K D+W+DD + +AD L Sbjct: 403 QCGLPYFEEWRKQMYMATSKNRALRPESYRDEWEDDDLVLQAQQDFADYL 452 >ref|XP_006307454.1| hypothetical protein CARUB_v10009080mg [Capsella rubella] gi|482576165|gb|EOA40352.1| hypothetical protein CARUB_v10009080mg [Capsella rubella] Length = 461 Score = 70.9 bits (172), Expect = 1e-09 Identities = 41/113 (36%), Positives = 54/113 (47%) Frame = +3 Query: 3 ELQSKWVARVLSGKVLLPTXXXXXXXXXXXXXXXXXAGMPKHYTHALPLIETKYKNCLLA 182 ELQSKWVA VLSG+V LP+ G PK YTH L +Y N + Sbjct: 338 ELQSKWVAAVLSGRVTLPSEEKMMEDVTADYAVRDSHGFPKRYTHRLGAGHVQYLNWIAE 397 Query: 183 EIGLPPLEEWREKLHEDTIKFIKSVQRGSSDQWDDDYWDAVIKSKYADLLANR 341 ++G PP EEWR + E + + D+WDDD+ +I Y D L + Sbjct: 398 QVGAPPGEEWRYQEIEGGYVRLATQSDTFRDKWDDDH---LILEAYEDFLRQK 447