BLASTX nr result
ID: Glycyrrhiza28_contig00008715
seq
BLASTX 2.2.26 [Sep-21-2011] Reference: Altschul, Stephen F., Thomas L. Madden, Alejandro A. Schaffer, Jinghui Zhang, Zheng Zhang, Webb Miller, and David J. Lipman (1997), "Gapped BLAST and PSI-BLAST: a new generation of protein database search programs", Nucleic Acids Res. 25:3389-3402. Query= Glycyrrhiza28_contig00008715 (559 letters) Database: ./nr 115,041,592 sequences; 42,171,959,267 total letters Searching..................................................done Score E Sequences producing significant alignments: (bits) Value XP_016189690.1 PREDICTED: autophagy-related protein 16 [Arachis ... 197 1e-57 KHN34375.1 Protein tipD [Glycine soja] 197 2e-57 XP_003525614.1 PREDICTED: protein tipD-like [Glycine max] KRH571... 197 2e-57 KYP65337.1 Protein tipD [Cajanus cajan] 196 2e-57 XP_015964550.1 PREDICTED: autophagy-related protein 16 [Arachis ... 196 3e-57 XP_004515850.1 PREDICTED: protein tipD [Cicer arietinum] 196 4e-57 XP_014625479.1 PREDICTED: protein tipD-like [Glycine max] KHN104... 195 1e-56 XP_019464314.1 PREDICTED: autophagy-related protein 16 [Lupinus ... 194 2e-56 GAU40407.1 hypothetical protein TSUD_136540 [Trifolium subterran... 193 5e-56 XP_007155503.1 hypothetical protein PHAVU_003G207100g [Phaseolus... 193 7e-56 XP_017972162.1 PREDICTED: autophagy-related protein 16 [Theobrom... 192 8e-56 XP_013460974.1 nucleotide-binding protein [Medicago truncatula] ... 192 1e-55 XP_017409583.1 PREDICTED: autophagy-related protein 16 isoform X... 192 2e-55 EOY22311.1 Transducin/WD40 repeat-like superfamily protein [Theo... 192 2e-55 XP_017409575.1 PREDICTED: autophagy-related protein 16 isoform X... 192 2e-55 XP_017409570.1 PREDICTED: autophagy-related protein 16 isoform X... 192 3e-55 XP_017409555.1 PREDICTED: autophagy-related protein 16 isoform X... 192 4e-55 XP_019457397.1 PREDICTED: autophagy-related protein 16-like [Lup... 191 5e-55 XP_010102923.1 Protein tipD [Morus notabilis] EXB94441.1 Protein... 190 9e-55 XP_010026927.1 PREDICTED: autophagy-related protein 16 isoform X... 189 2e-54 >XP_016189690.1 PREDICTED: autophagy-related protein 16 [Arachis ipaensis] XP_016189705.1 PREDICTED: autophagy-related protein 16 [Arachis ipaensis] Length = 514 Score = 197 bits (502), Expect = 1e-57 Identities = 88/101 (87%), Positives = 98/101 (97%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTLRA+GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLRASGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEH+SS+LCC WSG+GKPLASADKNG+VC+WT Sbjct: 474 SKGDIVSTLKEHSSSVLCCRWSGMGKPLASADKNGIVCVWT 514 >KHN34375.1 Protein tipD [Glycine soja] Length = 514 Score = 197 bits (501), Expect = 2e-57 Identities = 88/101 (87%), Positives = 97/101 (96%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTL+A GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLKAMGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEHTSS+LCC WSG+GKPLASADKNG+VC+WT Sbjct: 474 SKGDIVSTLKEHTSSVLCCRWSGIGKPLASADKNGIVCVWT 514 >XP_003525614.1 PREDICTED: protein tipD-like [Glycine max] KRH57171.1 hypothetical protein GLYMA_05G043700 [Glycine max] Length = 514 Score = 197 bits (501), Expect = 2e-57 Identities = 88/101 (87%), Positives = 97/101 (96%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTL+A GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLKAMGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEHTSS+LCC WSG+GKPLASADKNG+VC+WT Sbjct: 474 SKGDIVSTLKEHTSSVLCCRWSGIGKPLASADKNGIVCVWT 514 >KYP65337.1 Protein tipD [Cajanus cajan] Length = 487 Score = 196 bits (499), Expect = 2e-57 Identities = 87/101 (86%), Positives = 98/101 (97%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 +TSGRDNLHNLFDVRSLEV GTLRA+GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 387 VTSGRDNLHNLFDVRSLEVCGTLRASGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 446 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEH+SS+LCC WSG+GKPLASADKNG+VC+WT Sbjct: 447 SKGDIVSTLKEHSSSVLCCRWSGIGKPLASADKNGIVCVWT 487 >XP_015964550.1 PREDICTED: autophagy-related protein 16 [Arachis duranensis] XP_015964558.1 PREDICTED: autophagy-related protein 16 [Arachis duranensis] Length = 514 Score = 196 bits (499), Expect = 3e-57 Identities = 88/101 (87%), Positives = 97/101 (96%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTLRA+GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLRASGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEH+SS+LCC WSG+GKPLASADKNG VC+WT Sbjct: 474 SKGDIVSTLKEHSSSVLCCKWSGMGKPLASADKNGTVCVWT 514 >XP_004515850.1 PREDICTED: protein tipD [Cicer arietinum] Length = 511 Score = 196 bits (498), Expect = 4e-57 Identities = 87/101 (86%), Positives = 98/101 (97%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTLRATGNRVASNWSRSCISPD+NH+AAGSADGS+Y+WS Sbjct: 411 LTSGRDNLHNLFDVRSLEVCGTLRATGNRVASNWSRSCISPDNNHVAAGSADGSVYIWSI 470 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+G+IVSTLKEHTSS+LCC WSG+GKPLASAD+NG+VC+WT Sbjct: 471 SKGEIVSTLKEHTSSVLCCRWSGMGKPLASADRNGIVCVWT 511 >XP_014625479.1 PREDICTED: protein tipD-like [Glycine max] KHN10461.1 Protein tipD [Glycine soja] KRH03886.1 hypothetical protein GLYMA_17G126200 [Glycine max] KRH03887.1 hypothetical protein GLYMA_17G126200 [Glycine max] KRH03888.1 hypothetical protein GLYMA_17G126200 [Glycine max] KRH03889.1 hypothetical protein GLYMA_17G126200 [Glycine max] Length = 514 Score = 195 bits (495), Expect = 1e-56 Identities = 87/101 (86%), Positives = 96/101 (95%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTL+A GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLKAMGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEHTSS+LCC WSG+ KPLASADKNG+VC+WT Sbjct: 474 SKGDIVSTLKEHTSSVLCCRWSGIAKPLASADKNGIVCVWT 514 >XP_019464314.1 PREDICTED: autophagy-related protein 16 [Lupinus angustifolius] OIW00291.1 hypothetical protein TanjilG_27542 [Lupinus angustifolius] Length = 514 Score = 194 bits (494), Expect = 2e-56 Identities = 88/101 (87%), Positives = 96/101 (95%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTLRA GNRVASNWSRSCISPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLRAAGNRVASNWSRSCISPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEHTSS+L C WSG+GKPLASADKNG+VC+WT Sbjct: 474 SKGDIVSTLKEHTSSVLSCRWSGIGKPLASADKNGIVCLWT 514 >GAU40407.1 hypothetical protein TSUD_136540 [Trifolium subterraneum] Length = 514 Score = 193 bits (491), Expect = 5e-56 Identities = 89/101 (88%), Positives = 98/101 (97%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEVSGTL+A+GNRVASNWSRSCISP D ++AAGS+DGS+YVWS Sbjct: 414 LTSGRDNLHNLFDVRSLEVSGTLKASGNRVASNWSRSCISPGDKNVAAGSSDGSVYVWSV 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEHTSSILCC+WSGLG+PLASADKNGVVCIWT Sbjct: 474 SKGDIVSTLKEHTSSILCCSWSGLGRPLASADKNGVVCIWT 514 >XP_007155503.1 hypothetical protein PHAVU_003G207100g [Phaseolus vulgaris] XP_007155504.1 hypothetical protein PHAVU_003G207100g [Phaseolus vulgaris] ESW27497.1 hypothetical protein PHAVU_003G207100g [Phaseolus vulgaris] ESW27498.1 hypothetical protein PHAVU_003G207100g [Phaseolus vulgaris] Length = 514 Score = 193 bits (490), Expect = 7e-56 Identities = 87/101 (86%), Positives = 96/101 (95%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTL+ATGNRVASNWSRSCISPDDNH+AAG ADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLKATGNRVASNWSRSCISPDDNHVAAGYADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIV TLKEHTSS+L CTWSG+GKPLASADKNG+VC+WT Sbjct: 474 SKGDIVGTLKEHTSSVLGCTWSGVGKPLASADKNGIVCVWT 514 >XP_017972162.1 PREDICTED: autophagy-related protein 16 [Theobroma cacao] Length = 509 Score = 192 bits (489), Expect = 8e-56 Identities = 87/101 (86%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFD+RSLEV GT RA GNRVASNWSRSC+SPDDNHIAAGSADGSIY+WS Sbjct: 409 LTSGRDNLHNLFDIRSLEVCGTFRANGNRVASNWSRSCMSPDDNHIAAGSADGSIYIWSI 468 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+ DIVSTLKEHT+ +LCCTWSGLGKPLASADKNG+VC WT Sbjct: 469 SKADIVSTLKEHTAPVLCCTWSGLGKPLASADKNGIVCTWT 509 >XP_013460974.1 nucleotide-binding protein [Medicago truncatula] KEH35008.1 nucleotide-binding protein [Medicago truncatula] Length = 509 Score = 192 bits (488), Expect = 1e-55 Identities = 88/101 (87%), Positives = 96/101 (95%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFD+RSLEVSGTL+A GNRVASNWSRSCISPDD ++AAGS+DGS+YVWS Sbjct: 409 LTSGRDNLHNLFDMRSLEVSGTLKAAGNRVASNWSRSCISPDDKNVAAGSSDGSVYVWSA 468 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+ DIV TLKEHTSSILCCTWSGLG+PLASADKNGVVCIWT Sbjct: 469 SKADIVGTLKEHTSSILCCTWSGLGRPLASADKNGVVCIWT 509 >XP_017409583.1 PREDICTED: autophagy-related protein 16 isoform X4 [Vigna angularis] XP_017409589.1 PREDICTED: autophagy-related protein 16 isoform X4 [Vigna angularis] XP_017409596.1 PREDICTED: autophagy-related protein 16 isoform X4 [Vigna angularis] KOM32732.1 hypothetical protein LR48_Vigan01g228800 [Vigna angularis] Length = 514 Score = 192 bits (487), Expect = 2e-55 Identities = 86/101 (85%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV TL+A GNRVASNWSRSC SPDDNH+AAGSADGS+Y+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCSTLKAAGNRVASNWSRSCFSPDDNHVAAGSADGSVYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIV TLKEHTSS+LCCTWSG+ KPLASADKNGVVC+WT Sbjct: 474 SKGDIVGTLKEHTSSVLCCTWSGIRKPLASADKNGVVCVWT 514 >EOY22311.1 Transducin/WD40 repeat-like superfamily protein [Theobroma cacao] Length = 550 Score = 192 bits (489), Expect = 2e-55 Identities = 87/101 (86%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFD+RSLEV GT RA GNRVASNWSRSC+SPDDNHIAAGSADGSIY+WS Sbjct: 450 LTSGRDNLHNLFDIRSLEVCGTFRANGNRVASNWSRSCMSPDDNHIAAGSADGSIYIWSI 509 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+ DIVSTLKEHT+ +LCCTWSGLGKPLASADKNG+VC WT Sbjct: 510 SKADIVSTLKEHTAPVLCCTWSGLGKPLASADKNGIVCTWT 550 >XP_017409575.1 PREDICTED: autophagy-related protein 16 isoform X3 [Vigna angularis] Length = 528 Score = 192 bits (487), Expect = 2e-55 Identities = 86/101 (85%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV TL+A GNRVASNWSRSC SPDDNH+AAGSADGS+Y+WS Sbjct: 428 LTSGRDNLHNLFDVRSLEVCSTLKAAGNRVASNWSRSCFSPDDNHVAAGSADGSVYIWSI 487 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIV TLKEHTSS+LCCTWSG+ KPLASADKNGVVC+WT Sbjct: 488 SKGDIVGTLKEHTSSVLCCTWSGIRKPLASADKNGVVCVWT 528 >XP_017409570.1 PREDICTED: autophagy-related protein 16 isoform X2 [Vigna angularis] Length = 531 Score = 192 bits (487), Expect = 3e-55 Identities = 86/101 (85%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV TL+A GNRVASNWSRSC SPDDNH+AAGSADGS+Y+WS Sbjct: 431 LTSGRDNLHNLFDVRSLEVCSTLKAAGNRVASNWSRSCFSPDDNHVAAGSADGSVYIWSI 490 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIV TLKEHTSS+LCCTWSG+ KPLASADKNGVVC+WT Sbjct: 491 SKGDIVGTLKEHTSSVLCCTWSGIRKPLASADKNGVVCVWT 531 >XP_017409555.1 PREDICTED: autophagy-related protein 16 isoform X1 [Vigna angularis] XP_017409563.1 PREDICTED: autophagy-related protein 16 isoform X1 [Vigna angularis] BAT76005.1 hypothetical protein VIGAN_01395700 [Vigna angularis var. angularis] Length = 553 Score = 192 bits (487), Expect = 4e-55 Identities = 86/101 (85%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV TL+A GNRVASNWSRSC SPDDNH+AAGSADGS+Y+WS Sbjct: 453 LTSGRDNLHNLFDVRSLEVCSTLKAAGNRVASNWSRSCFSPDDNHVAAGSADGSVYIWSI 512 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIV TLKEHTSS+LCCTWSG+ KPLASADKNGVVC+WT Sbjct: 513 SKGDIVGTLKEHTSSVLCCTWSGIRKPLASADKNGVVCVWT 553 >XP_019457397.1 PREDICTED: autophagy-related protein 16-like [Lupinus angustifolius] XP_019457405.1 PREDICTED: autophagy-related protein 16-like [Lupinus angustifolius] OIW18309.1 hypothetical protein TanjilG_31449 [Lupinus angustifolius] Length = 514 Score = 191 bits (484), Expect = 5e-55 Identities = 88/101 (87%), Positives = 94/101 (93%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFDVRSLEV GTLRA GNRVASNWSRSCISPDDNH+AAGSADGSIY+WS Sbjct: 414 LTSGRDNLHNLFDVRSLEVCGTLRAAGNRVASNWSRSCISPDDNHVAAGSADGSIYIWSI 473 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S GDIVSTLKEHTSS+L C WSG+GK LASADKNG+VC+WT Sbjct: 474 STGDIVSTLKEHTSSVLSCRWSGIGKTLASADKNGIVCLWT 514 >XP_010102923.1 Protein tipD [Morus notabilis] EXB94441.1 Protein tipD [Morus notabilis] Length = 525 Score = 190 bits (483), Expect = 9e-55 Identities = 84/101 (83%), Positives = 96/101 (95%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDN+HNLFD+RSLEV GTLRATGNR+ASNWSRSC+SPDD+++AAGSADGS+Y+WS Sbjct: 425 LTSGRDNMHNLFDIRSLEVCGTLRATGNRIASNWSRSCLSPDDSYVAAGSADGSVYIWSI 484 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+ DIVSTLKEHTS +LCC+WSGLGKPLASADKNGVVC WT Sbjct: 485 SKADIVSTLKEHTSPVLCCSWSGLGKPLASADKNGVVCTWT 525 >XP_010026927.1 PREDICTED: autophagy-related protein 16 isoform X2 [Eucalyptus grandis] KCW83232.1 hypothetical protein EUGRSUZ_B00173 [Eucalyptus grandis] Length = 509 Score = 189 bits (480), Expect = 2e-54 Identities = 83/101 (82%), Positives = 96/101 (95%) Frame = -2 Query: 558 LTSGRDNLHNLFDVRSLEVSGTLRATGNRVASNWSRSCISPDDNHIAAGSADGSIYVWST 379 LTSGRDNLHNLFD+RSLEV GTLRATGNRVASNWSRSCISPDD+++AAGSADG++Y+WS Sbjct: 409 LTSGRDNLHNLFDMRSLEVCGTLRATGNRVASNWSRSCISPDDSYVAAGSADGAVYIWSI 468 Query: 378 SRGDIVSTLKEHTSSILCCTWSGLGKPLASADKNGVVCIWT 256 S+GDIVSTLKEHT+ +LCC+WSGLGKPL SADKNG++C WT Sbjct: 469 SKGDIVSTLKEHTAPVLCCSWSGLGKPLCSADKNGIICTWT 509